E. Woloszczak visited
the same region a decade later in the course of a systematic floristic
survey of the Eastern Carpathians. His studies covered the area between
the upper San and the Oslawa rivers and were published in 1894. These
provided the rationale for the proposed phytogeographical dividing line
between the Western and Eastern Carpathians from the £upkowska pass,
running north along the Oslawa and San rivers (Woloszczak 1908).
In
the period between two world wars, the area of Bieszczady did not
attract much interest from the naturalists, as the high summits of the
Czarnohora and Czywczynskie mountain ranges were far more interesting.
The interest in this area increased markedly in the early 1950s.
Systematic work started on forests, meadows and peat-bogs, and floristic
searches were conducted in various plant groups. It resulted in many
fundamental papers pertaining to mosses (Lisowski 1956), lichens (Glanc,
Tobolewski 1960), vascular plants (Jasiewicz 1965), and fungi (Domanski
1964, Domanski et al. 1960, 1963, 1967, 1970). The history of the
vegetation cover in this part of the Carpathians was studied by
Ralska-Jasiewiczowa (1980), whereas the issues of plant geography were
studied e.g. by Pawlowski (1972) and Zemanek (1991 a, b, 1992).
Detailed
studies on the flora of the Bieszczady National Park were carried out
from 1993–96. The inventory of vascular plants was performed in a grid
of squares 1 x 1 kilometre, using the methodology adopted for the in
Distribution atlas of vascular plants in Poland (ATPOL) (Zajac A., Zajac
M. [eds] 2001, Zemanek 1993). The results were included in an
unpublished report for the Park (Zemanek et al. MS.) and in a floristic
note reporting newly found taxa (Zemanek et al. 1996). A paper on the
whole flora of vascular plants in the Bieszczady National Park is
published in the “Bieszczady Monographs” (Zemanek, Winnicki 1999).
The
results of the floristic studies of the 1950s and 1960s, documenting
the great wealth and uniqueness of the Bieszczady flora were used as
arguments to support the cause for establishing the Park in 1973.
The
vascular plants are the best studied group of plants in the Bieszczady
National Park. In the whole Western Bieszczady ca. 850 were noted while
in the Polish part of the Eastern Carpathians ca. 1100 species (Zemanek
1991a). At present, ca. 780 species are reported to occur in the Park
and this number increases with intensified studies. The species added to
the list are mainly those associated with the activities of Man (weeds,
ruderal plants). Some findings were unexpected and surprising, such as
the finding in 1996 of a station with alpine larkspur Delphinum elatum, a
subalpine species, whose nearest stations are located at Bor¿awa, some
60 kilometres to the south-east.
Apart
from new findings there are some species which disappeared. As a result
of irresponsible collectors’ activities it is highly probable that an
extremely rare long-flowered primrose Primula longiflora (=P. hallerii)
that once occurred in the area has now disappeared for ever. Several
species are seriously threatened due to intensive tourist traffic and
the trampling on the summit regions of the Tarnica and Halicz summits,
the rocky crests of the Smerek, Polonina Caryñska, Polonina Wetliñska
and Rozsypaniec. These are locations with the most valuable components
of high mountain flora.
The
taxa representing the eastern element confirm the classification of the
Bieszczady mountains within the Eastern Carpathians and are the unique
feature which distinguish the Bieszczady National Park from all other
Polish national parks. There are 30 such taxa, including 6 Eastern
Carpathian endemic species: white cow-wheat Melampyrum saxosum —
occurring in the subalpine zone of the Kinczyk Bukowski and on the
Polonina Bukowska summits, Eastern Carpathian monkshood Aconitum
lasiocarpum, fairly common throughout the area, dwarf monkshood A.
bucovinense, one of the rarest plants in Poland, occurring on only four
stations and not exceeding several dozen in number, Carthusian pink
Dianthus carthusianorum subsp. saxigenus, occurring on the rocks of the
Krzemieñ and Bukowe Berdo summits, and probably, Turkul lady’s mantle
Alchemilla turkulensis, found in tall-herb communities on the poloninas.
Among the interesting East-Carpathian plant species, the following can
be found on poloninas: Kotschy’s knapweed Centaurea kotschyana, Eastern
Carpathian thistle Cirsium waldsteinii, compact pink Dianthus compactus,
rose vipergrass Scorzonera rosea, and Dacian violet Viola dacica. In
forests, scopolia Scopolia carniolica and stoloniferous bellflower
Campanula abietina can be found, while in spring the alder woods upon
the Terebowiec and Wo³osaty brooks show white carpets of the flowering
spring snowflake Leucoium vernum subsp. carpaticum.
Most
of the taxa classified in the eastern element reach the limits of their
distribution ranges in the Park or in the Bieszczady mountains as such,
only a few also occur further west in the Western Carpathians.
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